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HMBR is an analogue with an additional methyl group on the aromatic ring and is non-fluorescent. HMBR conjugated with Y-FAST emits yellow fluorescence under blue light excitation (Ex= 419 nm; Em= 525–539 nm). HMBR is non-toxic to zebrafish embryos. HMBR has high cell permeability[1].

[1] Plamont MA, et al. Small fluorescence-activating and absorption-shifting tag for tunable protein imaging in vivo [published correction appears in Proc Natl Acad Sci U S A. 2016 Mar 8;113(10):E1412. Moncoq, Karine [added]]. Proc Natl Acad Sci U S A. 2016;113(3):497-502. DOI:10.1073/pnas.1513094113
[2] Zhenping Cao, et al. Encoding with a fluorescence-activating and absorption-shifting tag generates living bacterial probes for mammalian microbiota imaging. Mater Today Bio. 2022 Jun 6:15:100311. DOI:10.1016/j.mtbio.2022.100311